1946
Born in Brooklyn, New York, June 19
1963
Received New York State Regents Scholarship
1963
Entered College of Agriculture, Cornell University
1964
Appointed part-time Assistant to Curator, Insect Collections, Cornell University
1964
Member of New York (Brooklyn) Entomological Society
1965
Member of Cornell University Mexico Field Research Party (with Dr. W. L. Brown, Jr.)
1965-1968
Ford Three-Year Scholar's Program
1966
Appointed Field Research Assistant to Dr. Thomas Eisner at Archbold Biological Station in Florida and at Cave Creek Ranch, Portal, Arizona
1966-1968
Part-time Assistant to Librarian, Entomology Library, Cornell University
1967
Appointed Field Research Assistant (with D. Simberloff for Dr. E. O. Wilson) in Florida Keys and Arizona
1967
Received B.S., Cornell University
1967
Entered Cornell University, Department of Biology as graduate student
1968
Received M.S., Cornell University
1968
Entered Harvard University as post-graduate
1968-1973
Teaching fellowship, Department of Biology, Harvard University
1968-1982
Member of Cambridge Entomological Society
1969
Field work at Archbold Biological Station in Florida
1969
Elected Vice President, Cambridge Entomological Society
1969-1973
Richmond Fellowship, Harvard University
1970
Member of Graduate Student Council, Department of Biology, Harvard University
1970
Elected President, Cambridge Entomological Society
1970
Field work at Charles Darwin Research Station in the Galapagos Islands
1970
Field work at the E. N. Huyck Preserve, Albany County, New York
1971
Field work in Florida, Arizona, and Colorado
1972
Participated in Organization for Tropical Studies trip to Costa Rica
1972
Field work in Nova Scotia, Kansas, Colorado, and Arizona
1973
Received Ph.D., Harvard University
1973
Field work in Costa Rica, Guatemala, and Ecuador
1973
Published "Ultraviolet Differences Between Sulphur Butterflies"
1973-1974
Member of Executive Committee, Cambridge Entomological Society
1973-1977
Member of Library Committee, Museum of Comparative Zoology, Harvard University
1973-1978
Assistant Professor of Biology, Harvard University
1973-1978
Assistant Curator of Lepidoptera, Museum of Comparative Zoology, Harvard University
1973-1982
Consultant to the Biological Laboratories Insectary Facility, Harvard University
1973-1982
Member of Lignum vitae Key (Florida) University Council
1973-1982
Member of Editorial Board, Psyche
1974
Field work at Archbold Biological Station in Florida
1974-1975
Section Vice Chairman, Section A (Systematics, Morphology and Evolution), Entomological Society of America
1974-1975
Member of Scanning Electron Microscope Committee, Museum of Comparative Zoology, Harvard University
1974-1980
Member of Scanning Electron Microscope Facility Committee, Department of Biology, Harvard University 1974-1981 member of Entomological Society of America
1974-1980
Member of Governing Board, Organization for Tropical Studies (representing Harvard University)
1975
Participated in a safari to Ethiopia, Kenya, and Tanzania
1975
Served on Program Evaluation Committee, Section A (Systematics, Morphology and Evolution), Entomological Society of America
1975-1976
Section Chairman, Section A (Systematics, Morphology and Evolution), Entomological Society of America
1975-1980
Contributing reviewer for American Reference Book Annual
1976
Appointed Biologist, Smithsonian Tropical Research Institute (STRI)
1976
Member of Nominating Committee and Special Constitution and By-laws Revision Committee, Section A (Systematics, Morphology and Evolution), Entomological Society of America
1976-1977
Organized and administered Natural History Seminar series of Department of Biology, Harvard University
1976-1980
Member of Education Committee, Organization for Tropical Studies
1977
Field work in Panama, Ecuador, and Colombia
1977
Published "Communication in the Lepidoptera"
1977
Member of Committee on Meeting Dates, Section A (Systematics, Morphology and Evolution), Entomological Society of America
1977-1980
Member of Executive Committee, Organization for Tropical Studies
1977-1981
Contributing reviewer forLibrary Journal
1977-1982
Staff Scientist (Research Entomologist), STRI
1978
Field work in the Dominican Republic published "Ultraviolet Reflection and its Behavioral Role in the Courtship of the Sulfur Butterflies Colias eurytheme and C. philodice"
1978
Contributing reviewer for Reprint Bulletin, Book Reviews
1978-1981
Associate Professor of Biology, Harvard University
1978-1981
Hessel Associate Curator of Lepidoptera, Museum of Comparative Zoology, Harvard University
1978-1981
Scientist-in-Charge, Barro Colorado Island research station, STRI
1979
Published "Communications in the Ultraviolet"
1980
Field work in Colombia
1981
Guest speaker at the Royal Entomological Society International Symposium on the Biology of Butterflies, Reading, England
1981-1982
Member of Executive Council, Lepidopterists' Society
1982
Died in Washington, D.C., January 13
1984
Posthumously published "Visual Communication and Sexual Selection Among Butterflies"